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  1. Wellesley was born at Harley Street, Marylebone, London, the eldest son of Arthur Wellesley, 1st Duke of Wellington, and the Honourable Catherine Sarah Dorothea "Kitty" Pakenham, daughter of Edward Pakenham, 2nd Baron Longford. Lord Charles Wellesley was his younger brother and Lord Wellesley, Lord Mornington and Lord Cowley his uncles.

  2. Valerian Wellesley, 8th Duke of Wellington (2 July 1915 – 31 December 2014) Lady Elizabeth Wellesley (26 December 1918 – 25 November 2013) Dorothy, a poet, became the lover of Vita Sackville-West, (who wrote her entry for the Oxford Dictionary of National Biography). Wellesley had been engaged to Sackville-West's lover Violet Trefusis.